I would expect that the A4000T has the ROMs in the same chip positions (U175/176). Most of the 4000T motherboard is common with the 4000D.
Of course, the main difference between them is that the 4000T had real SCSI on board (like the 3000). But I don't think that would stop it booting altogether, just make some error messages or crashes.
My bet is your ROMs are screwed.
tony