If the gurus with both expansions are the same (or very similar) the mainboard and/or the Agnus socket seems to be the problem.
You'll need to open the case and take a look at the trapdoor connector - possibly some of the acid damage has spread over there and some cleaning/resoldering is required. Additionally, the Agnus socket can become unreliable with old age and improper storage. I may have to pull out the Agnus chip to inspect/clean the contacts - make sure you use a PLCC puller (~3€). Levering with a screwdriver can very quickly break the socket, especially with age.
Ah yes: the RAM address lines come from Agnus (it's the RAM controller for chip RAM). The data lines run through some latches (U10-U13) to the CPU side. (The latch is the point where the busses can be deconnected to run independently: chipset side and CPU/fast side.)